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Abstract Atmospheric rivers (ARs) reaching high-latitudes in summer contribute to the majority of climatological poleward water vapor transport into the Arctic. This transport has exhibited long term changes over the past decades, which cannot be entirely explained by anthropogenic forcing according to ensemble model responses. Here, through observational analyses and model experiments in which winds are adjusted to match observations, we demonstrate that low-frequency, large-scale circulation changes in the Arctic play a decisive role in regulating AR activity and thus inducing the recent upsurge of this activity in the region. It is estimated that the trend in summertime AR activity may contribute to 36% of the increasing trend of atmospheric summer moisture over the entire Arctic since 1979 and account for over half of the humidity trends in certain areas experiencing significant recent warming, such as western Greenland, northern Europe, and eastern Siberia. This indicates that AR activity, mostly driven by strong synoptic weather systems often regarded as stochastic, may serve as a vital mechanism in regulating long term moisture variability in the Arctic. 

Abstract Increasing severity of extreme heat is a hallmark of climate change. Its impacts depend on temperature but also on moisture and solar radiation, each with distinct spatial patterns and vertical profiles. Here, we consider these variables’ combined effect on extreme heat stress, as measured by the environmental stress index, using a suite of high-resolution climate simulations for historical (1980–2005) and future (2074–2099, Representative Concentration Pathway 8.5 (RCP8.5)) periods. We find that observed extreme heat stress drops off nearly linearly with elevation above a coastal zone, at a rate that is larger in more humid regions. Future projections indicate dramatic relative increases whereby the historical top 1% summer heat stress value may occur on about 25%–50% of future summer days under the RCP8.5 scenario. Heat stress increases tend to be larger at higher latitudes and in areas of greater temperature increase, although in the southern and eastern US moisture increases are nearly as important. Imprinted on top of this dominant pattern we find secondary effects of smaller heat stress increases near ocean coastlines, notably along the Pacific coast, and larger increases in mountains, notably the Sierra Nevada and southern Appalachians. This differential warming is attributable to the greater warming of land relative to ocean, and to larger temperature increases at higher elevations outweighing larger water-vapor increases at lower elevations. All together, our results aid in furthering knowledge about drivers and characteristics that shape future extreme heat stress at scales difficult to capture in global assessments. 

Atmospheric rivers (ARs) are long, narrow synoptic scale weather features important for Earth’s hydrological cycle typically transporting water vapor poleward, delivering precipitation important for local climates. Understanding ARs in a warming climate is problematic because the AR response to climate change is tied to how the feature is defined. The Atmospheric River Tracking Method Intercomparison Project (ARTMIP) provides insights into this problem by comparing 16 atmospheric river detection tools (ARDTs) to a common data set consisting of high resolution climate change simulations from a global atmospheric general circulation model. ARDTs mostly show increases in frequency and intensity, but the scale of the response is largely dependent on algorithmic criteria. Across ARDTs, bulk characteristics suggest intensity and spatial footprint are inversely correlated, and most focus regions experience increases in precipitation volume coming from extreme ARs. The spread of the AR precipitation response under climate change is large and dependent on ARDT selection. 

Abstract Statistical relationships between atmospheric rivers (ARs) and extratropical cyclones and anticyclones are investigated on a global scale using objectively identified ARs, cyclones, and anticyclones during 1979–2014. Composites of circulation and moisture fields around the ARs show that a strong cyclone is located poleward and westward of the AR centroid, which confirms the close link between the AR and extratropical cyclone. In addition, a pronounced anticyclone is found to be located equatorward and eastward of the AR, whose presence together with the cyclone leads to strong horizontal pressure gradient that forces moisture to be transported along a narrow corridor within the warm sector of the cyclone. This anticyclone located toward the downstream equatorward side of the cyclone is found to be missing for cyclones not associated with ARs. These key features are robust in composites performed in different hemispheres, over different ocean basins, and with respect to different AR intensities. Furthermore, correlation analysis shows that the AR intensity is much better correlated with the pressure gradient between the cyclone and anticyclone than with the cyclone/anticyclone intensity alone, although stronger cyclones favor the occurrence of AR. The importance of the horizontal pressure gradient in the formation of the AR is also consistent with the fact that climatologically ARs are frequently found over the region between the polar lows and subtropical highs in all seasons. 

Abstract Atmospheric rivers (ARs) are filaments of enhanced horizontal moisture transport in the atmosphere. Due to their prominent role in the meridional moisture transport and regional weather extremes, ARs have been studied extensively in recent years. Yet, the representations of ARs and their associated precipitation on a global scale remains largely unknown. In this study, we developed an AR detection algorithm specifically for satellite observations using moisture and the geostrophic winds derived from 3D geopotential height field from the combined retrievals of the Atmospheric Infrared Sounder and the Advanced Microwave Sounding Unit on NASA Aqua satellite. This algorithm enables us to develop the first global AR catalog based solely on satellite observations. The satellite‐based AR catalog is then combined with the satellite‐based precipitation (Integrated Muti‐SatellitE Retrievals for GPM) to evaluate the representations of ARs and AR‐induced precipitation in reanalysis products. Our results show that the spreads in AR frequency and AR length distribution are generally small across data sets, while the spread in AR width is relatively larger. Reanalysis products are found to consistently underestimate both mean and extreme AR‐related precipitation. However, all reanalyses tend to precipitate too often under AR conditions, especially over low latitude regions. This finding is consistent with the “drizzling” bias which has plagued generations of climate models. Overall, the findings of this study can help to improve the representations of ARs and associated precipitation in reanalyses and climate models. 

How homeodomain proteins gain sufficient specificity to control different cell fates has been a long-standing problem in developmental biology. The conserved Gsx homeodomain proteins regulate specific aspects of neural development in animals from flies to mammals, and yet they belong to a large transcription factor family that bind nearly identical DNA sequences in vitro. Here, we show that the mouse and fly Gsx factors unexpectedly gain DNA binding specificity by forming cooperative homodimers on precisely spaced and oriented DNA sites. High-resolution genomic binding assays revealed that Gsx2 binds both monomer and homodimer sites in the developing mouse ventral telencephalon. Importantly, reporter assays showed that Gsx2 mediates opposing outcomes in a DNA binding site-dependent manner: Monomer Gsx2 binding represses transcription, whereas homodimer binding stimulates gene expression. In Drosophila , the Gsx homolog, Ind, similarly represses or stimulates transcription in a site-dependent manner via an autoregulatory enhancer containing a combination of monomer and homodimer sites. Integrating these findings, we test a model showing how the homodimer to monomer site ratio and the Gsx protein levels defines gene up-regulation versus down-regulation. Altogether, these data serve as a new paradigm for how cooperative homeodomain transcription factor binding can increase target specificity and alter regulatory outcomes. 

Abstract The atmospheric river (AR) frequency trends over the Southern Hemisphere are investigated using three reanalyses and two Community Earth System Model (CESM) ensembles. The results show that AR frequency has been increasing over the Southern Ocean and decreasing over lower latitudes in the past four decades and that ARs have been shifting poleward. While the observed trends are mostly driven by the poleward shift of the westerly jet, fully coupled CESM experiments indicate anthropogenic forcing would result in positive AR frequency trends over the Southern Ocean due mostly to moisture changes. The difference between the observed trends and anthropogenically driven trends can be largely reconciled by the atmosphere‐only CESM simulations forced by observed sea surface temperatures: Sea surface temperature variability characteristic of the negative phase of the Interdecadal Pacific Oscillation strongly suppresses the moisture‐driven trends while enhances the circulation‐induced trends over the Southern Ocean, thus bringing the simulated trends into closer agreement with the observed trends.